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Research suggests elevated MMP-1 (a collagen-degrading enzyme) may contribute

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In both UC and CD, 25% of the samples showed a significant increase in C

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Other reported mechanisms include activation of FAK-paxillin complexes (which help cells migrate and attach to surfaces during wound healing), JAK-2 signaling, and ERK1/2 pathway activity (Sikiric et al., Current Neuropharmacology, 2016