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Although the molecular mechanisms underpinning anhydrobiosis are not yet completely understood, it seems clear that desiccation tolerance depends on the synergic action of an array of many different molecules working together

The carbonyl placement in the oxyanion hole likely positions the electrophilic center in the proper orientation for nucleophilic attack by Cys95 and explains why alternative warheads are not useful for this scaffold against UCHL3

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5), indicating that in the latter case zinc remained bound to the protein as it would otherwise exert cytotoxic action (Supplementary Fig