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Subsequently, the membranes were incubated with secondary antibodies for another 3 h

To put it simply, skin aging is a combination of intrinsic factors (like your natural genetic clock and collagen depletion) and extrinsic factors (like sun damage, pollution, and facial expressions)

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B.SakK.et al (2019)

Panno ML, Giordano F, Rizza P, Pellegrino M, Zito D, Giordano C, Mauro L, Catalano S, Aquila S, Sisci D, De Amicis F, Vivacqua A, Fuqua SW, And S (2012) Bergapten induces ER depletion in breast cancer cells through SMAD4-mediated ubiquitination